diff --git a/lib/jasmine-core/jasmine.js b/lib/jasmine-core/jasmine.js
index e79fe8f2..b7188584 100644
--- a/lib/jasmine-core/jasmine.js
+++ b/lib/jasmine-core/jasmine.js
@@ -1490,12 +1490,18 @@ getJasmineRequireObj().Env = function(j$) {
     };
 
     var makeMatchersUtil = function() {
-      var customEqualityTesters =
-        runnableResources[currentRunnable().id].customEqualityTesters;
-      return new j$.MatchersUtil({
-        customTesters: customEqualityTesters,
-        pp: makePrettyPrinter()
-      });
+      const cr = currentRunnable();
+
+      if (cr) {
+        const customEqualityTesters =
+          runnableResources[cr.id].customEqualityTesters;
+        return new j$.MatchersUtil({
+          customTesters: customEqualityTesters,
+          pp: makePrettyPrinter()
+        });
+      } else {
+        return new j$.MatchersUtil({ pp: j$.basicPrettyPrinter_ });
+      }
     };
 
     var expectationFactory = function(actual, spec) {
@@ -2026,7 +2032,8 @@ getJasmineRequireObj().Env = function(j$) {
         }
 
         return undefined;
-      }
+      },
+      makeMatchersUtil
     );
 
     var spyRegistry = new j$.SpyRegistry({
@@ -8647,11 +8654,6 @@ getJasmineRequireObj().Spy = function(j$) {
     };
   })();
 
-  var matchersUtil = new j$.MatchersUtil({
-    customTesters: [],
-    pp: j$.makePrettyPrinter()
-  });
-
   /**
    * @classdesc _Note:_ Do not construct this directly. Use {@link spyOn},
    * {@link spyOnProperty}, {@link jasmine.createSpy}, or
@@ -8659,19 +8661,24 @@ getJasmineRequireObj().Spy = function(j$) {
    * @class Spy
    * @hideconstructor
    */
-  function Spy(name, originalFn, customStrategies, defaultStrategyFn) {
+  function Spy(name, matchersUtil, optionals) {
+    const { originalFn, customStrategies, defaultStrategyFn } = optionals || {};
+
     var numArgs = typeof originalFn === 'function' ? originalFn.length : 0,
       wrapper = makeFunc(numArgs, function(context, args, invokeNew) {
         return spy(context, args, invokeNew);
       }),
-      strategyDispatcher = new SpyStrategyDispatcher({
-        name: name,
-        fn: originalFn,
-        getSpy: function() {
-          return wrapper;
+      strategyDispatcher = new SpyStrategyDispatcher(
+        {
+          name: name,
+          fn: originalFn,
+          getSpy: function() {
+            return wrapper;
+          },
+          customStrategies: customStrategies
         },
-        customStrategies: customStrategies
-      }),
+        matchersUtil
+      ),
       callTracker = new j$.CallTracker(),
       spy = function(context, args, invokeNew) {
         /**
@@ -8783,11 +8790,11 @@ getJasmineRequireObj().Spy = function(j$) {
     return wrapper;
   }
 
-  function SpyStrategyDispatcher(strategyArgs) {
+  function SpyStrategyDispatcher(strategyArgs, matchersUtil) {
     var baseStrategy = new j$.SpyStrategy(strategyArgs);
     var argsStrategies = new StrategyDict(function() {
       return new j$.SpyStrategy(strategyArgs);
-    });
+    }, matchersUtil);
 
     this.and = baseStrategy;
 
@@ -8816,9 +8823,10 @@ getJasmineRequireObj().Spy = function(j$) {
     };
   }
 
-  function StrategyDict(strategyFactory) {
+  function StrategyDict(strategyFactory, matchersUtil) {
     this.strategies = [];
     this.strategyFactory = strategyFactory;
+    this.matchersUtil = matchersUtil;
   }
 
   StrategyDict.prototype.any = function() {
@@ -8843,7 +8851,7 @@ getJasmineRequireObj().Spy = function(j$) {
     var i;
 
     for (i = 0; i < this.strategies.length; i++) {
-      if (matchersUtil.equals(args, this.strategies[i].args)) {
+      if (this.matchersUtil.equals(args, this.strategies[i].args)) {
         return this.strategies[i].strategy;
       }
     }
@@ -8853,16 +8861,19 @@ getJasmineRequireObj().Spy = function(j$) {
 };
 
 getJasmineRequireObj().SpyFactory = function(j$) {
-  function SpyFactory(getCustomStrategies, getDefaultStrategyFn) {
+  function SpyFactory(
+    getCustomStrategies,
+    getDefaultStrategyFn,
+    getMatchersUtil
+  ) {
     var self = this;
 
     this.createSpy = function(name, originalFn) {
-      return j$.Spy(
-        name,
+      return j$.Spy(name, getMatchersUtil(), {
         originalFn,
-        getCustomStrategies(),
-        getDefaultStrategyFn()
-      );
+        customStrategies: getCustomStrategies(),
+        defaultStrategyFn: getDefaultStrategyFn()
+      });
     };
 
     this.createSpyObj = function(baseName, methodNames, propertyNames) {
diff --git a/spec/core/SpySpec.js b/spec/core/SpySpec.js
index 6271c900..bfc08e98 100644
--- a/spec/core/SpySpec.js
+++ b/spec/core/SpySpec.js
@@ -231,7 +231,7 @@ describe('Spies', function() {
     expect(spy('baz', 'grault', 'waldo')).toEqual(42);
   });
 
-  it('uses custom equality testers when selecting a strategy', function() {
+  it('uses asymmetric equality testers when selecting a strategy', function() {
     var spy = env.createSpy('foo');
     spy.and.returnValue(42);
     spy.withArgs(jasmineUnderTest.any(String)).and.returnValue(-1);
@@ -240,6 +240,23 @@ describe('Spies', function() {
     expect(spy({})).toEqual(42);
   });
 
+  it('uses the provided matchersUtil selecting a strategy', function() {
+    const matchersUtil = new jasmineUnderTest.MatchersUtil({
+      customTesters: [
+        function(a, b) {
+          if ((a === 'bar' && b === 'baz') || (a === 'baz' && b === 'bar')) {
+            return true;
+          }
+        }
+      ]
+    });
+    const spy = new jasmineUnderTest.Spy('aSpy', matchersUtil);
+    spy.and.returnValue('default strategy return value');
+    spy.withArgs('bar').and.returnValue('custom strategy return value');
+    expect(spy('foo')).toEqual('default strategy return value');
+    expect(spy('baz')).toEqual('custom strategy return value');
+  });
+
   it('can reconfigure an argument-specific strategy', function() {
     var spy = env.createSpy('foo');
     spy.withArgs('foo').and.returnValue(42);
diff --git a/spec/core/integration/EnvSpec.js b/spec/core/integration/EnvSpec.js
index 462893d9..2d0b0072 100644
--- a/spec/core/integration/EnvSpec.js
+++ b/spec/core/integration/EnvSpec.js
@@ -3413,4 +3413,56 @@ describe('Env integration', function() {
       done();
     });
   });
+
+  it('uses custom equality testers in Spy#withArgs', async function() {
+    env.it('a spec', function() {
+      const createSpySpy = env.createSpy('via createSpy');
+      const spiedOn = { foo: function() {} };
+      env.spyOn(spiedOn, 'foo');
+      const spyObj = env.createSpyObj('spyObj', ['foo']);
+      const spiedOnAllFuncs = { foo: function() {} };
+      env.spyOnAllFunctions(spiedOnAllFuncs);
+
+      for (spy of [
+        createSpySpy,
+        spiedOn.foo,
+        spyObj.foo,
+        spiedOnAllFuncs.foo
+      ]) {
+        spy.and.returnValue('default strategy');
+        spy.withArgs(42).and.returnValue('custom strategy');
+      }
+
+      env.addCustomEqualityTester(function(a, b) {
+        if ((a === 'x' && b === 42) || (a === 42 && b === 'x')) {
+          return true;
+        }
+      });
+
+      env
+        .expect(createSpySpy('x'))
+        .withContext('createSpy')
+        .toEqual('custom strategy');
+      env
+        .expect(spiedOn.foo('x'))
+        .withContext('spyOn')
+        .toEqual('custom strategy');
+      env
+        .expect(spyObj.foo('x'))
+        .withContext('createSpyObj')
+        .toEqual('custom strategy');
+      env
+        .expect(spiedOnAllFuncs.foo('x'))
+        .withContext('spyOnAllFunctions')
+        .toEqual('custom strategy');
+    });
+
+    let failedExpectations;
+    env.addReporter({
+      specDone: r => (failedExpectations = r.failedExpectations)
+    });
+
+    await env.execute();
+    expect(failedExpectations).toEqual([]);
+  });
 });
diff --git a/src/core/Env.js b/src/core/Env.js
index 4ee1eae1..ddd9d4ee 100644
--- a/src/core/Env.js
+++ b/src/core/Env.js
@@ -327,12 +327,18 @@ getJasmineRequireObj().Env = function(j$) {
     };
 
     var makeMatchersUtil = function() {
-      var customEqualityTesters =
-        runnableResources[currentRunnable().id].customEqualityTesters;
-      return new j$.MatchersUtil({
-        customTesters: customEqualityTesters,
-        pp: makePrettyPrinter()
-      });
+      const cr = currentRunnable();
+
+      if (cr) {
+        const customEqualityTesters =
+          runnableResources[cr.id].customEqualityTesters;
+        return new j$.MatchersUtil({
+          customTesters: customEqualityTesters,
+          pp: makePrettyPrinter()
+        });
+      } else {
+        return new j$.MatchersUtil({ pp: j$.basicPrettyPrinter_ });
+      }
     };
 
     var expectationFactory = function(actual, spec) {
@@ -863,7 +869,8 @@ getJasmineRequireObj().Env = function(j$) {
         }
 
         return undefined;
-      }
+      },
+      makeMatchersUtil
     );
 
     var spyRegistry = new j$.SpyRegistry({
diff --git a/src/core/Spy.js b/src/core/Spy.js
index 7b7e422b..26d4b6b4 100644
--- a/src/core/Spy.js
+++ b/src/core/Spy.js
@@ -7,11 +7,6 @@ getJasmineRequireObj().Spy = function(j$) {
     };
   })();
 
-  var matchersUtil = new j$.MatchersUtil({
-    customTesters: [],
-    pp: j$.makePrettyPrinter()
-  });
-
   /**
    * @classdesc _Note:_ Do not construct this directly. Use {@link spyOn},
    * {@link spyOnProperty}, {@link jasmine.createSpy}, or
@@ -19,19 +14,24 @@ getJasmineRequireObj().Spy = function(j$) {
    * @class Spy
    * @hideconstructor
    */
-  function Spy(name, originalFn, customStrategies, defaultStrategyFn) {
+  function Spy(name, matchersUtil, optionals) {
+    const { originalFn, customStrategies, defaultStrategyFn } = optionals || {};
+
     var numArgs = typeof originalFn === 'function' ? originalFn.length : 0,
       wrapper = makeFunc(numArgs, function(context, args, invokeNew) {
         return spy(context, args, invokeNew);
       }),
-      strategyDispatcher = new SpyStrategyDispatcher({
-        name: name,
-        fn: originalFn,
-        getSpy: function() {
-          return wrapper;
+      strategyDispatcher = new SpyStrategyDispatcher(
+        {
+          name: name,
+          fn: originalFn,
+          getSpy: function() {
+            return wrapper;
+          },
+          customStrategies: customStrategies
         },
-        customStrategies: customStrategies
-      }),
+        matchersUtil
+      ),
       callTracker = new j$.CallTracker(),
       spy = function(context, args, invokeNew) {
         /**
@@ -143,11 +143,11 @@ getJasmineRequireObj().Spy = function(j$) {
     return wrapper;
   }
 
-  function SpyStrategyDispatcher(strategyArgs) {
+  function SpyStrategyDispatcher(strategyArgs, matchersUtil) {
     var baseStrategy = new j$.SpyStrategy(strategyArgs);
     var argsStrategies = new StrategyDict(function() {
       return new j$.SpyStrategy(strategyArgs);
-    });
+    }, matchersUtil);
 
     this.and = baseStrategy;
 
@@ -176,9 +176,10 @@ getJasmineRequireObj().Spy = function(j$) {
     };
   }
 
-  function StrategyDict(strategyFactory) {
+  function StrategyDict(strategyFactory, matchersUtil) {
     this.strategies = [];
     this.strategyFactory = strategyFactory;
+    this.matchersUtil = matchersUtil;
   }
 
   StrategyDict.prototype.any = function() {
@@ -203,7 +204,7 @@ getJasmineRequireObj().Spy = function(j$) {
     var i;
 
     for (i = 0; i < this.strategies.length; i++) {
-      if (matchersUtil.equals(args, this.strategies[i].args)) {
+      if (this.matchersUtil.equals(args, this.strategies[i].args)) {
         return this.strategies[i].strategy;
       }
     }
diff --git a/src/core/SpyFactory.js b/src/core/SpyFactory.js
index 808ec647..03bd6035 100644
--- a/src/core/SpyFactory.js
+++ b/src/core/SpyFactory.js
@@ -1,14 +1,17 @@
 getJasmineRequireObj().SpyFactory = function(j$) {
-  function SpyFactory(getCustomStrategies, getDefaultStrategyFn) {
+  function SpyFactory(
+    getCustomStrategies,
+    getDefaultStrategyFn,
+    getMatchersUtil
+  ) {
     var self = this;
 
     this.createSpy = function(name, originalFn) {
-      return j$.Spy(
-        name,
+      return j$.Spy(name, getMatchersUtil(), {
         originalFn,
-        getCustomStrategies(),
-        getDefaultStrategyFn()
-      );
+        customStrategies: getCustomStrategies(),
+        defaultStrategyFn: getDefaultStrategyFn()
+      });
     };
 
     this.createSpyObj = function(baseName, methodNames, propertyNames) {